Table 1 Genetic variation and relatedness using nine microsatellite loci in nine colonies. For colony r, the standard error (SE) was jackknifed over loci. For mean r, colonies were weighted equally and SE was jackknifed over groups. A = alleles per locus per colony, He = binomial expected heterozygosity, r = relatedness.

r (SE) Colony A He All Females 1 7.0 0.747 0.1650.055 0.2000.068 2 5.6 0.777 0.0910.023 0.0950.026 3 9.0 0.834 0.0310.016 0.0370.022 4 8.6 0.791 0.1200.017 0.1310.017 5 5.2 0.741 0.1920.055 0.2400.074 6 5.2 0.754 0.1590.032 0.1720.036 7 8.3 0.794 0.1150.032 0.1180.031 8 4.4 0.691 0.3070.051 0.3480.058 9 5.9 0.819 0.0730.033 0.0740.043 Mean 6.6 0.775 0.1360.023 0.1530.028