Figure 1. The annual colony cycle of Lasioglossum (Chilalictus) hemichalceum. In spring, colonies consist of mated, overwintered females (blue) some females forage early (light blue) and others do not forage until later (dark blue). Foragers die before their brood matures but unrelated adults are present in colonies because not all adults forage at the same time. The hatched vertical line indicates timing of this study.